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<DIV><A href="http://www3.interscience.wiley.com/journal/117963685/home"><FONT 
color=#000000>Journal of Biogeography</FONT></A></DIV>
<DIV><A href="http://www3.interscience.wiley.com/journal/123534348/issue"><FONT 
color=#000000>Volume 37 Issue 
7</FONT></A>,&nbsp;Pages&nbsp;1352&nbsp;-&nbsp;1362</DIV>
<DIV class=pubOnline>Published Online: 23&nbsp;Mar&nbsp;2010</DIV>
<DIV class=title-document xmlns:jencode="java.net.URLEncoder">&nbsp;</DIV>
<DIV class=title-document xmlns:jencode="java.net.URLEncoder"><FONT 
size=4>Biogeographical contingency and the evolution of tropical anchovies 
(genus <SPAN class=i><EM>Cetengraulis</EM></SPAN>) from temperate anchovies 
(genus <SPAN class=i><EM>Engraulis</EM></SPAN>)</FONT></DIV>
<DIV class=author-info xmlns="http://www.w3.org/1999/xhtml" 
xmlns:jencode="java.net.URLEncoder"><SPAN class=name><SPAN class=forenames>W. 
S.</SPAN> <SPAN class=surname>Grant</SPAN> <SUP><NOBR>1</NOBR></SUP>*</SPAN>, 
<SPAN class=name><SPAN class=forenames>Frédéric</SPAN> <SPAN 
class=surname>Lecomte</SPAN> <SUP><NOBR>2</NOBR></SUP> </SPAN>and <SPAN 
class=name><SPAN class=forenames>Brian W.</SPAN> <SPAN 
class=surname>Bowen</SPAN> <SUP><NOBR>3</NOBR></SUP> </SPAN></DIV>
<DIV class=addresses xmlns="http://www.w3.org/1999/xhtml" 
xmlns:jencode="java.net.URLEncoder"><SPAN class=address><A 
class=invisible-anchor name=a1>&nbsp;</A><SPAN id=a1> <SPAN 
class=number><SUP><NOBR>1</NOBR></SUP> </SPAN>Department of Biological Sciences, 
University of Alaska Anchorage, Anchorage, AK 99508, USA and Alaska Department 
of Fish and Game, 333 Raspberry Road, Anchorage, AK 99518, USA</SPAN> , <A 
class=invisible-anchor name=a2>&nbsp;</A><SPAN id=a2> <SPAN 
class=number><SUP><NOBR>2</NOBR></SUP> </SPAN>INRA-UMR ECOBIOP, Pôle 
d'Hydrobiologie, Quartier Ibarron, 64310 Saint Pée sur Nivelle, France</SPAN> , 
<A class=invisible-anchor name=a3>&nbsp;</A><SPAN id=a3> <SPAN 
class=number><SUP><NOBR>3</NOBR></SUP> </SPAN>Hawaii Institute of Marine 
Biology, University of Hawaii, PO Box 1346, Kaneohe, HI, 96744, 
USA</SPAN>&nbsp;</SPAN><A class=invisible-anchor name=c1>&nbsp;</A><SPAN id=c1> 
<SPAN class=number>*</SPAN>Correspondence: Stewart Grant, Department of 
Biological Sciences, University of Alaska Anchorage, Anchorage, AK 99508, USA. 
E-mail: <A class=externallink href="mailto:phylogeo@yahoo.com"><FONT 
color=#000000>phylogeo@yahoo.com</FONT></A></SPAN> </DIV>
<DIV class=bpg40copyright>&nbsp;</DIV>
<DIV class=abstract-title id=h1><STRONG>ABSTRACT</STRONG></DIV>
<DIV class=abstract-content xmlns="http://www.w3.org/1999/xhtml" 
xmlns:jencode="java.net.URLEncoder" xmlns:md="urn:mdransfi-functions" 
xmlns:f="http://wiley.com/wispers/content/function" 
xmlns:xs="http://www.w3.org/2001/XMLSchema">
<P class=para><SPAN class=h5-inline><STRONG>Aim&nbsp;</STRONG></SPAN>Similar 
regimes of selection in different geographical settings can deterministically 
produce similar adaptive morphologies. We tested the hypothesis that the 
evolutionary trajectories of fish in upwelling zones can be altered by 
biogeographic contingencies in the biological and physical environment.</P>
<P class=para><SPAN 
class=h5-inline><STRONG>Location&nbsp;</STRONG></SPAN>Eastern Pacific and 
western Atlantic oceans.</P>
<P class=para><SPAN class=h5-inline><STRONG>Methods&nbsp;</STRONG></SPAN>We 
estimated phylogenetic relationships among eastern Pacific temperate anchovies 
(genus <SPAN class=i><EM>Engraulis</EM></SPAN>) and tropical anchovies (genus 
<SPAN class=i><EM>Cetengraulis</EM></SPAN>) with neighbour-joining and Bayesian 
tree analysis of a 521-bp segment of mitochondrial DNA cytochrome <SPAN 
class=i><EM>b</EM></SPAN>. Available sequences for five additional engraulid 
taxa were included to establish polarity of the tree. Bayesian estimates (BEAST) 
of time to most recent common ancestor (TMRCA) for the nodes in the phylogeny 
were calibrated with divergence between <SPAN class=i><EM>Cetengraulis 
edentulus</EM></SPAN> and <SPAN class=i><EM>Cetengraulis mysticetus</EM></SPAN> 
precipitated by the rise of the Panama Isthmus 2.8–3.2&nbsp;Ma.</P>
<P class=para><SPAN 
class=h5-inline><STRONG>Results&nbsp;</STRONG></SPAN>Neighbour-joining and 
Bayesian trees indicate that<U> South American <SPAN class=i><EM>Engraulis 
anchoita</EM></SPAN> (Argentina) and <SPAN class=i><EM>Engraulis 
ringens</EM></SPAN> (Chile) together are basal sister taxa to the California 
anchovy (<SPAN class=i><EM>Engraulis mordax</EM></SPAN>) and Old World anchovies 
(<SPAN class=i><EM>Engraulis japonicas</EM></SPAN>, <SPAN class=i><EM>Engraulis 
australis</EM></SPAN>, <SPAN class=i><EM>Engraulis capensis</EM></SPAN> and 
<SPAN class=i><EM>Engraulis encrasicolus</EM></SPAN>)</U>. The two tropical 
species of <SPAN class=i><EM>Cetengraulis</EM></SPAN> are sister-taxa to 
Californian <SPAN class=i><EM>E. mordax</EM></SPAN>, even though their 
phenotypes and ecologies differ markedly. A relaxed molecular clock indicates a 
TMRCA between Californian <SPAN class=i><EM>E. mordax</EM></SPAN> and <SPAN 
class=i><EM>Cetengraulis</EM></SPAN> at about 4.2&nbsp;Ma (3.0–6.3&nbsp;Ma 95% 
highest probability density).</P>
<P class=para><SPAN class=h5-inline><STRONG>Main 
conclusions&nbsp;</STRONG></SPAN>The TMRCA between the California anchovy, <SPAN 
class=i><EM>E. mordax</EM></SPAN>, and tropical <SPAN 
class=i><EM>Cetengraulis</EM></SPAN> coincides with the formation of the Gulf of 
California, which provided opportunities for allopatric isolation during climate 
oscillations. Mid-Pliocene warming (3.1–2.9&nbsp;Ma) may have trapped ancestors 
of <SPAN class=i><EM>Cetengraulis</EM></SPAN> in the Gulf of California, where 
they evolved digestive tract morphologies to exploit inshore tropical habitats 
with low plankton productivities. While populations of several other temperate 
fishes have become isolated in the Gulf of California, few of these derived 
species show strong adaptive shifts from temperate sister taxa or range 
expansions into the tropical provinces of the western Atlantic and eastern 
Pacific.</P></DIV></FONT></BODY></HTML>