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<div class=Section1>

<p class=MsoNormal style='background:white;vertical-align:baseline'><cite><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333300;
border:none windowtext 1.0pt;padding:0in;font-style:normal'>Science</span></cite><span
class=apple-converted-space><span style='font-size:12.0pt;font-family:"Arial","sans-serif";
color:#333300;border:none windowtext 1.0pt;padding:0in'> </span></span><span
class=slug-pub-date><span style='font-size:12.0pt;font-family:"Arial","sans-serif";
color:#333300;border:none windowtext 1.0pt;padding:0in'>28 November 2014:</span></span><span
class=apple-converted-space><span style='font-size:12.0pt;font-family:"Arial","sans-serif";
color:#333300;border:none windowtext 1.0pt;padding:0in'> </span></span><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333300;
border:none windowtext 1.0pt;padding:0in'><br>
<span class=slug-vol>Vol. 346</span><span class=apple-converted-space> </span><span
class=slug-issue>no. 6213</span><span class=apple-converted-space> </span><span
class=slug-pages>pp. 1102-1106</span><span class=apple-converted-space> </span><br>
<cite><span style='font-family:"Arial","sans-serif";font-style:normal'>DOI:</span></cite><span
class=apple-converted-space> </span><span class=slug-doi>10.1126/science.1258396</span></span><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333300'><o:p></o:p></span></p>

<h1 style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:1.5pt;
margin-left:0in;line-height:12.5pt;background:white;vertical-align:baseline'><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333333'><o:p> </o:p></span></h1>

<h1 style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:1.5pt;
margin-left:0in;mso-line-height-alt:12.5pt;background:white;vertical-align:
baseline'><span style='font-size:16.0pt;font-family:"Arial","sans-serif";
color:#333333'>Increasing anthropogenic nitrogen in the North Pacific Ocean<o:p></o:p></span></h1>

<p class=MsoNormal style='margin-left:0in;text-indent:-.25in;mso-list:l0 level1 lfo2;
background:white;vertical-align:baseline'><![if !supportLists]><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333333'><span
style='mso-list:Ignore'>1.<span style='font:7.0pt "Times New Roman"'>   
</span></span></span><![endif]><span class=name><span style='font-size:12.0pt;
font-family:"Arial","sans-serif";color:#333333;border:none windowtext 1.0pt;
padding:0in'><a
href="http://www.sciencemag.org/search?author1=Il-Nam+Kim&sortspec=date&submit=Submit"><span
style='color:#333333'>Il-Nam Kim</span></a></span></span><span
style='font-size:12.0pt;font-family:"Arial","sans-serif";color:#333333'><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-1"
id=xref-aff-1-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>1</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=Kitack+Lee&sortspec=date&submit=Submit"><span
style='color:#333333'>Kitack Lee</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-1"
id=xref-aff-1-2><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>1</span></sup></a><span class=xref-sep><sup><span
style='border:none windowtext 1.0pt;padding:0in'>,</span></sup></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#corresp-1"
id=xref-corresp-1-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>*</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=Nicolas+Gruber&sortspec=date&submit=Submit"><span
style='color:#333333'>Nicolas Gruber</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-2"
id=xref-aff-2-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>2</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=David+M.+Karl&sortspec=date&submit=Submit"><span
style='color:#333333'>David M. Karl</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-3"
id=xref-aff-3-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>3</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=John+L.+Bullister&sortspec=date&submit=Submit"><span
style='color:#333333'>John L. Bullister</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-4"
id=xref-aff-4-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>4</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=Simon+Yang&sortspec=date&submit=Submit"><span
style='color:#333333'>Simon Yang</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-2"
id=xref-aff-2-2><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>2</span></sup></a>,<span class=apple-converted-space> </span><span
class=name><span style='border:none windowtext 1.0pt;padding:0in'><a
href="http://www.sciencemag.org/search?author1=Tae-Wook+Kim&sortspec=date&submit=Submit"><span
style='color:#333333'>Tae-Wook Kim</span></a></span></span><a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#aff-5"
id=xref-aff-5-1><sup><span style='color:#2E6D8F;border:none windowtext 1.0pt;
padding:0in'>5</span></sup></a><o:p></o:p></span></p>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l3 level1 lfo3;
background:white;vertical-align:baseline'><a name=aff-1></a><![if !supportLists]><span
style='font-family:"Arial","sans-serif";color:#333333;font-style:normal'><span
style='mso-list:Ignore'>1.<span style='font:7.0pt "Times New Roman"'>   
</span></span></span><![endif]><sup><span style='font-family:"Arial","sans-serif";
color:#333333;border:none windowtext 1.0pt;padding:0in;font-style:normal'>1</span></sup><span
style='font-family:"Arial","sans-serif";color:#333333;font-style:normal'>School
of Environmental Sciences and Engineering, Pohang University of Science and
Technology (POSTECH), Pohang, 790–784, Republic of Korea.<o:p></o:p></span></address>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l3 level1 lfo3;
background:white;vertical-align:baseline;outline-style: none;font-weight:inherit;
font-size:inherit;line-height:inherit;text-align:inherit'><a name=aff-2
id=aff-2></a><![if !supportLists]><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'><span style='mso-list:Ignore'>2.<span
style='font:7.0pt "Times New Roman"'>    </span></span></span><![endif]><sup><span
style='font-family:"Arial","sans-serif";color:#333333;border:none windowtext 1.0pt;
padding:0in;font-style:normal'>2</span></sup><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'>Environmental Physics Group, Institute of
Biogeochemistry and Pollutant Dynamics, ETH Zürich, Zürich, Switzerland.<o:p></o:p></span></address>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l3 level1 lfo3;
background:white;vertical-align:baseline;outline-style: none;font-weight:inherit;
font-size:inherit;line-height:inherit;text-align:inherit'><a name=aff-3
id=aff-3></a><![if !supportLists]><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'><span style='mso-list:Ignore'>3.<span
style='font:7.0pt "Times New Roman"'>    </span></span></span><![endif]><sup><span
style='font-family:"Arial","sans-serif";color:#333333;border:none windowtext 1.0pt;
padding:0in;font-style:normal'>3</span></sup><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'>Daniel K. Inouye Center for Microbial
Oceanography, University of Hawaii at Manoa, 1950 East West Road, Honolulu, HI
96822, USA.<o:p></o:p></span></address>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l3 level1 lfo3;
background:white;vertical-align:baseline;outline-style: none;font-weight:inherit;
font-size:inherit;line-height:inherit;text-align:inherit'><a name=aff-4
id=aff-4></a><![if !supportLists]><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'><span style='mso-list:Ignore'>4.<span
style='font:7.0pt "Times New Roman"'>    </span></span></span><![endif]><sup><span
style='font-family:"Arial","sans-serif";color:#333333;border:none windowtext 1.0pt;
padding:0in;font-style:normal'>4</span></sup><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'>Pacific Marine Environmental Laboratory,
National Oceanic and Atmospheric Administration (NOAA), Seattle, WA 98115, USA.<o:p></o:p></span></address>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l2 level1 lfo4;
background:white;vertical-align:baseline;outline-style: none;font-weight:inherit;
font-size:inherit;line-height:inherit;text-align:inherit'><a name=aff-5
id=aff-5></a><![if !supportLists]><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'><span style='mso-list:Ignore'>1.<span
style='font:7.0pt "Times New Roman"'>    </span></span></span><![endif]><sup><span
style='font-family:"Arial","sans-serif";color:#333333;border:none windowtext 1.0pt;
padding:0in'>5</span></sup><span style='font-family:"Arial","sans-serif";
color:#333333'>Ocean Circulation and Climate Research Division, Korea Institute
of Ocean Science and Technology, Ansan, 426–744, Republic of Korea.<a
href="http://www.sciencemag.org/content/346/6213/1102.abstract?utm_campaign=email-sci-toc&utm_source=eloqua#xref-corresp-1-1"><span
style='font-family:"Lucida Sans Unicode","sans-serif";color:#2E6D8F;border:
none windowtext 1.0pt;padding:0in'>↵</span></a><span class=corresp-label><sup><span
style='border:none windowtext 1.0pt;padding:0in'>*</span></sup></span>Corresponding
author. E-mail:</span><span class=apple-converted-space><span style='font-family:
"Arial","sans-serif";color:#333333'> </span></span><span style='font-family:
"Arial","sans-serif";color:#333333'><a href="mailto:ktl@postech.ac.kr"><span
style='color:#2E6D8F;border:none windowtext 1.0pt;padding:0in'>ktl@postech.ac.kr</span></a></span><span
style='font-family:"Arial","sans-serif";color:#333333;font-style:normal'><o:p></o:p></span></address>

<address style='margin-left:0in;text-indent:-.25in;mso-list:l2 level1 lfo4;
background:white;vertical-align:baseline'><![if !supportLists]><span
style='font-family:"Arial","sans-serif";color:#333333;font-style:normal'><span
style='mso-list:Ignore'>2.<span style='font:7.0pt "Times New Roman"'>   
</span></span></span><![endif]><span style='font-family:"Arial","sans-serif";
color:#333333;font-style:normal'><o:p> </o:p></span></address>

<p class=MsoNormal style='margin-left:0in;text-indent:-.25in;line-height:12.5pt;
mso-list:l1 level1 lfo5;background:white;vertical-align:baseline'><![if !supportLists]><span
style='font-size:10.0pt;font-family:Symbol;color:#333333'><span
style='mso-list:Ignore'>·<span style='font:7.0pt "Times New Roman"'>        
</span></span></span><![endif]><b><span style='font-size:12.0pt;font-family:
"Arial","sans-serif";color:#333333'><a
href="http://www.sciencemag.org/content/346/6213/tab-abstract"><span
style='text-transform:uppercase;border:solid #DADADA 1.0pt;padding:0in;
background:#EFEFEF'>ABSTRACT</span></a><o:p></o:p></span></b></p>

<p style='mso-margin-top-alt:12.0pt;margin-right:0in;margin-bottom:12.0pt;
margin-left:0in;background:#EEEEEE;vertical-align:baseline'><b><span
style='font-family:"Arial","sans-serif";color:#333333'>The recent increase in
anthropogenic emissions of reactive nitrogen from northeastern Asia and the
subsequent enhanced deposition over the extensive regions of the North Pacific
Ocean (NPO) have led to a detectable increase in the nitrate (N) concentration
of the upper ocean. The rate of increase of excess N relative to phosphate (P) was
found to be highest (</span></b><b><span style='font-family:"Cambria Math","serif";
color:#333333'>∼</span></b><b><span style='font-family:"Arial","sans-serif";
color:#333333'>0.24 micromoles per kilogram per year) in the vicinity of the
Asian source continent, with rates decreasing eastward across the NPO,
consistent with the magnitude and distribution of atmospheric nitrogen
deposition. This anthropogenically driven increase in the N content of the
upper NPO may enhance primary production in this N-limited region, potentially
leading to a long-term change of the NPO from being N-limited to P-limited.<o:p></o:p></span></b></p>

<p class=MsoNormal><o:p> </o:p></p>

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