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<div class=Section1>
<p class=MsoNormal><i><span style='font-family:"Arial","sans-serif"'>Nature </span></i><b><span
style='font-family:"Arial","sans-serif"'>543</span></b><span style='font-family:
"Arial","sans-serif"'>, 51–59<o:p></o:p></span></p>
<p class=MsoNormal><span style='font-family:"Arial","sans-serif"'>(02 March
2017)<o:p></o:p></span></p>
<p class=MsoNormal><span style='font-family:"Arial","sans-serif"'>doi:10.1038/nature21058<o:p></o:p></span></p>
<p class=MsoNormal><span style='font-family:"Arial","sans-serif"'>Published
online 01 March 2017<o:p></o:p></span></p>
<h1 style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:15.0pt;
margin-left:0in'><span style='font-size:8.0pt;font-family:"Arial","sans-serif";
color:#222222;letter-spacing:-.4pt;font-weight:normal'><o:p> </o:p></span></h1>
<h1 style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:15.0pt;
margin-left:0in'><span style='font-size:21.5pt;font-family:"Arial","sans-serif";
color:#222222;letter-spacing:-.4pt;font-weight:normal'>The integral role of
iron in ocean biogeochemistry<o:p></o:p></span></h1>
<p class=MsoNormal style='margin-left:0in;text-indent:-.25in;mso-list:l5 level1 lfo1'><![if !supportLists]><span
style='font-size:10.0pt;font-family:Symbol'><span style='mso-list:Ignore'>·<span
style='font:7.0pt "Times New Roman"'>
</span></span></span><![endif]><span style='font-family:"Arial","sans-serif"'><a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-1"><span
class=fn><span style='color:windowtext;text-decoration:none'>Alessandro
Tagliabue</span></span></a><span class=comma>,</span> <a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-2"><span
class=fn><span style='color:windowtext;text-decoration:none'>Andrew R. Bowie</span></span></a><span
class=comma>,</span> <a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-3"><span
class=fn><span style='color:windowtext;text-decoration:none'>Philip W. Boyd</span></span></a><span
class=comma>,</span> <a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-4"><span
class=fn><span style='color:windowtext;text-decoration:none'>Kristen N. Buck</span></span></a><span
class=comma>,</span> <a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-5"><span
class=fn><span style='color:windowtext;text-decoration:none'>Kenneth S. Johnson</span></span></a>
&<span class=apple-converted-space> </span><a
href="http://www.nature.com/nature/journal/v543/n7643/full/nature21058.html?WT.ec_id=NATURE-20170302&spMailingID=53531555&spUserID=MjA1NTE2ODQxMAS2&spJobID=1120165995&spReportId=MTEyMDE2NTk5NQS2#auth-6"><span
class=fn><span style='color:windowtext;text-decoration:none'>Mak A. Saito</span></span></a><o:p></o:p></span></p>
<h1 style='mso-margin-top-alt:13.5pt;margin-right:0in;margin-bottom:0in;
margin-left:0in;margin-bottom:.0001pt'><span style='font-size:14.0pt;
font-family:"Arial","sans-serif";color:#444444;letter-spacing:-.4pt'>Abstract<o:p></o:p></span></h1>
<p style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:15.0pt;
margin-left:0in'><span style='font-family:"Arial","sans-serif"'>The
micronutrient iron is now recognized to be important in regulating the
magnitude and dynamics of ocean primary productivity, making it an integral
component of the ocean’s biogeochemical cycles. In this Review, we
discuss how a recent increase in observational data for this trace metal has
challenged the prevailing view of the ocean iron cycle. Instead of focusing on
dust as the major iron source and emphasizing iron’s tight biogeochemical
coupling to major nutrients, a more complex and diverse picture of the sources
of iron, its cycling processes and intricate linkages with the ocean carbon and
nitrogen cycles has emerged.<o:p></o:p></span></p>
<h3 style='margin:0in;margin-bottom:.0001pt;background:whitesmoke'><span
style='font-size:11.0pt;color:#707070'><a href="javascript:;"><span
style='color:#363636;border:solid #9EABB4 1.0pt;padding:10.0pt;text-decoration:
none'>Figures</span></a><o:p></o:p></span></h3>
<p class=MsoNormal align=center style='text-align:center;line-height:14.4pt;
background:whitesmoke'><span style='font-size:9.0pt;color:#939393'><a
href="javascript:;"><span style='color:#939393;text-decoration:none'>First</span></a><span
class=apple-converted-space> </span>|<span class=apple-converted-space> </span></span><span
class=item-count><span style='font-size:9.0pt;color:#333333'>1-3 of 4</span></span><span
class=apple-converted-space><span style='font-size:9.0pt;color:#939393'> </span></span><span
style='font-size:9.0pt;color:#939393'>|<span class=apple-converted-space> </span><a
href="javascript:;"><span style='color:#5C7996;text-decoration:none'>Last</span></a><o:p></o:p></span></p>
<p class=MsoNormal style='background:whitesmoke'><a href="javascript:;"><b><span
style='color:#5C7996;border:none windowtext 1.0pt;padding:0in;text-decoration:
none'>left</span></b></a><span style='font-size:12.0pt'><o:p></o:p></span></p>
<p class=MsoNormal style='margin-left:0in;text-indent:-.25in;mso-list:l4 level1 lfo6;
background:whitesmoke'><![if !supportLists]><span style='mso-list:Ignore'>1.<span
style='font:7.0pt "Times New Roman"'> </span></span><![endif]><span
style='border:solid #CCCCCC 1.0pt;padding:0in;background:white'><a
href="http://www.nature.com/nature/journal/v543/n7643/fig_tab/nature21058_F1.html"><span
style='color:#5C7996;text-decoration:none'><img border=0 width=227 height=200
id="Imagen_x0020_1" src="cid:image001.jpg@01D2967A.780621D0"
alt="A schematic representation of the first view of the processes governing the ocean iron cycle."></span></a></span><span
class=legend><span style='font-size:9.0pt;color:#006699'>Figure 1</span></span><o:p></o:p></p>
<p class=MsoNormal style='margin-left:7.5pt;text-indent:-.25in;mso-list:l4 level1 lfo6;
background:whitesmoke'><![if !supportLists]><span style='mso-list:Ignore'>2.<span
style='font:7.0pt "Times New Roman"'> </span></span><![endif]><span
style='border:solid #CCCCCC 1.0pt;padding:0in;background:white'><a
href="http://www.nature.com/nature/journal/v543/n7643/fig_tab/nature21058_F2.html"><span
style='color:#5C7996;text-decoration:none'><img border=0 width=416 height=200
id="Imagen_x0020_2" src="cid:image002.jpg@01D2967A.780621D0"
alt="A revised representation of the major processes in the ocean iron cycle, with emphasis on the Atlantic Ocean."></span></a></span><span
class=legend><span style='font-size:9.0pt;color:#006699'>Figure 2</span></span><o:p></o:p></p>
<p class=MsoNormal style='margin-left:7.5pt;text-indent:-.25in;mso-list:l4 level1 lfo6;
background:whitesmoke'><![if !supportLists]><span style='mso-list:Ignore'>3.<span
style='font:7.0pt "Times New Roman"'> </span></span><![endif]><span
class=legend><span style='font-size:9.0pt;color:#006699'>Figure 3</span></span><o:p></o:p></p>
<p class=MsoNormal style='margin-left:7.5pt;text-indent:-.25in;mso-list:l4 level1 lfo6;
background:whitesmoke'><![if !supportLists]><span style='mso-list:Ignore'>4.<span
style='font:7.0pt "Times New Roman"'> </span></span><![endif]><span
style='border:solid #CCCCCC 1.0pt;padding:0in;background:white'><a
href="http://www.nature.com/nature/journal/v543/n7643/fig_tab/nature21058_F4.html"><span
style='color:#5C7996;text-decoration:none'><img border=0 width=682 height=200
id="Imagen_x0020_4" src="cid:image003.jpg@01D2967A.780621D0"
alt="Processes controlling the oceanic distribution of iron."></span></a></span><span
class=legend><span style='font-size:9.0pt;color:#006699'>Figure 4</span></span><o:p></o:p></p>
<p class=MsoNormal style='background:whitesmoke'><a href="javascript:;"><b><span
style='color:#5C7996;border:none windowtext 1.0pt;padding:0in;text-decoration:
none'>right</span></b></a><o:p></o:p></p>
<h2 style='margin:0in;margin-bottom:.0001pt;line-height:16.8pt;background:white'><span
style='font-size:9.0pt;text-transform:uppercase'>READ THE FULL ARTICLE<o:p></o:p></span></h2>
<p class=MsoNormal><o:p> </o:p></p>
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