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<p class=MsoNormal><span style='font-family:"Arial","sans-serif"'>Marine
Ecology</span><span style='font-family:"Arial","sans-serif"'><o:p></o:p></span></p>

<p class=MsoNormal style='background:white'><span class=epub-state><span
style='font-family:"Arial","sans-serif"'>First published: </span></span><span
class=epub-date><span style='font-family:"Arial","sans-serif"'>04 September 2018</span></span><span
style='font-family:"Arial","sans-serif"'><o:p></o:p></span></p>

<p class=MsoNormal style='line-height:14.7pt;background:white'><span
style='font-family:"Arial","sans-serif"'><a
href="https://doi.org/10.1111/maec.12503"><span style='color:windowtext'>https://doi.org/10.1111/maec.12503</span></a><o:p></o:p></span></p>

<h2 style='mso-margin-top-alt:7.5pt;margin-right:0in;margin-bottom:11.25pt;
margin-left:0in;line-height:22.5pt;background:white'><span style='font-family:
"Arial","sans-serif";color:#1C1D1E'><o:p> </o:p></span></h2>

<h2 style='mso-margin-top-alt:7.5pt;margin-right:0in;margin-bottom:11.25pt;
margin-left:0in;line-height:22.5pt;background:white'><span style='font-family:
"Arial","sans-serif";color:#1C1D1E'>Tidal height and sand as potential drivers
of the ecological interaction of the two intertidal mussels <i>Perumytilus
purpuratus</i><span class=apple-converted-space> </span>and<span
class=apple-converted-space> </span><i>Semimytilus algosus</i><o:p></o:p></span></h2>

<p class=MsoNormal style='line-height:14.7pt;background:white'><span
style='font-size:10.5pt;font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Pinochet%2C+Javier"
id="a1_Ctrl"><span lang=ES style='color:windowtext'>Javier Pinochet</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Dom%C3%ADnguez%2C+Joaqu%C3%ADn"><span
lang=ES style='color:windowtext'>Joaquín Domínguez</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Neira%2C+Evelyn"><span
lang=ES style='color:windowtext'>Evelyn Neira</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Rojas%2C+Camila"><span
lang=ES style='color:windowtext'>Camila Rojas</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Acu%C3%B1a%2C+Esteban"><span
lang=ES style='color:windowtext'>Esteban Acuña</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Cancino%2C+Juan+M"><span
lang=ES style='color:windowtext'>Juan M. Cancino</span></a></span><span
class=apple-converted-space><span lang=ES style='font-size:10.5pt;font-family:
"Arial","sans-serif"'> </span></span><span style='font-size:10.5pt;
font-family:"Arial","sans-serif"'><a
href="https://onlinelibrary.wiley.com/action/doSearch?ContribAuthorStored=Brante%2C+Antonio"><span
lang=ES style='color:windowtext'>Antonio Brante</span></a></span><span lang=ES
style='font-size:10.5pt;font-family:"Arial","sans-serif"'><o:p></o:p></span></p>

<p class=MsoNormal style='line-height:14.7pt;background:white'><span lang=ES
style='font-size:10.5pt;font-family:"Arial","sans-serif";color:#8B8B8B'><o:p> </o:p></span></p>

<h3 style='mso-margin-top-alt:0in;margin-right:0in;margin-bottom:11.25pt;
margin-left:0in;background:white'><span style='font-size:16.0pt;font-family:
"Arial","sans-serif";color:#414141'>Abstract<o:p></o:p></span></h3>

<p style='mso-margin-top-alt:3.75pt;margin-right:0in;margin-bottom:12.0pt;
margin-left:0in;background:white;box-sizing: border-box'><span
style='font-size:10.5pt;font-family:"Arial","sans-serif";color:#1C1D1E'>The
mussels<span class=apple-converted-space> </span><i>Perumytilus purpuratus</i><span
class=apple-converted-space> </span>and<span class=apple-converted-space> </span><i>Semimytilus
algosus</i><span class=apple-converted-space> </span>are two dominant
species of intertidal rocky shores of central Chile. These species have marked
differences in their distribution patterns with<span
class=apple-converted-space> </span><i>P. purpuratus</i><span
class=apple-converted-space> </span>dominating the mid</span><span
style='font-size:10.5pt;font-family:"Cambria Math","serif";color:#1C1D1E'>‐</span><span
style='font-size:10.5pt;font-family:"Arial","sans-serif";color:#1C1D1E'>intertidal
zone and<span class=apple-converted-space> </span><i>S. algosus</i><span
class=apple-converted-space> </span>dominating the lower intertidal zone
usually in habitats influenced by sand. Although it has been suggested that
differences in tolerance to environmental conditions, such as air exposure and
presence of sand, can explain the distribution of these species, there are
currently no experimental studies to support such hypotheses. Here, we
evaluated the growth and survival rate in the field of both mussel species at
four different tidal heights: 25, 75, 135, and 175 cm above the zero tide.
In addition, filtration rates were estimated for both species in the presence
and absence of sand in laboratory conditions. The results showed that shell and
wet weight growth rates of<span class=apple-converted-space> </span><i>P. purpuratus</i><span
class=apple-converted-space> </span>were highest in mid</span><span
style='font-size:10.5pt;font-family:"Cambria Math","serif";color:#1C1D1E'>‐</span><span
style='font-size:10.5pt;font-family:"Arial","sans-serif";color:#1C1D1E'> and
medium</span><span style='font-size:10.5pt;font-family:"Cambria Math","serif";
color:#1C1D1E'>‐</span><span style='font-size:10.5pt;font-family:"Arial","sans-serif";
color:#1C1D1E'>high tidal heights, whereas the growth rate of<span
class=apple-converted-space> </span><i>S. algosus</i><span
class=apple-converted-space> </span>was highest in the medium</span><span
style='font-size:10.5pt;font-family:"Cambria Math","serif";color:#1C1D1E'>‐</span><span
style='font-size:10.5pt;font-family:"Arial","sans-serif";color:#1C1D1E'>low
level. Similar pattern was observed for survival percentage. Furthermore, small<span
class=apple-converted-space> </span><i>S. algosus</i><span
class=apple-converted-space> </span>individuals cultivated with sand in
suspension had significantly higher filtration rates than<span
class=apple-converted-space> </span><i>P. purpuratus</i>. In large
individuals, no differences were observed between the two species nor between
treatments (presence and absence of sand). These results indicate that the
differences in the distribution patterns of<span class=apple-converted-space> </span><i>P. purpuratus</i><span
class=apple-converted-space> </span>and<span class=apple-converted-space> </span><i>S. algosus</i><span
class=apple-converted-space> </span>in the intertidal can be explained by
differences in physiological tolerances to both air exposure and to the
presence of sand.<o:p></o:p></span></p>

<p class=MsoNormal><o:p> </o:p></p>

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